This is a really well thought out alternative playstyle for Thrive. Thanks for making this!
I want to comment on one specific thing:
While the storage cost is as you say: you need slightly more. The other part I don’t think is really true. I think that most players play mobile builds, i.e. they want to move around. So most players already play with the movement energy cost being basically always on. Thus I’d say that a purely from balancing to normal gameplay standpoint, I would have set the osmoregulation multiplier not so low.